four ssp or forms of H. melpomene fly in Sangay NP : H. m. plessen, H. m. ecuadorensis, H. m. malletti and H. m. plesseni f. mimetica,
they're clearly supposed to be parapatric, but, in one area South of Puyo, they all fly together, giving birth to all kinds of mixed forms.
we present a sample of these forms, by "pattern" ; here, on the individual variation page, pattern "ecuadorensis with no yellow pigment".